Polydesmida: Paradoxosomatidae: Australiosomatinae:
Australiosomatini: Heterocladosoma species 


Heterocladosoma asperum (L. Koch, 1867)

Heterocladosoma asperum
Strongylosoma aspera L. Koch 1867. [From secondary source, reference not yet checked]
Orthomorpha aspera Attems 1898b: 328 (new combination)
Strongylosoma asperum Attems 1914: 235 (in species catalogue)
Orthomorpha aspera Chamberlin 1920a: 113 (in species catalogue)
Strongylosoma asperum Jeekel 1968: 19 (in list of Australian species)
   27 (in family revision, as species of 'enigmatic' generic status)
'Strongylosoma asperum' Jeekel 1981: 49 (in list of species recorded from eastern Australia)
Heterocladosoma asperum Jeekel 1987: 12 (in list of species recorded from Qld; new combination)
   24 (in species key)
   28 (redescription)
   Fig. 9 (p. 30)
 
Types: At least one male, Brisbane, Qld, date and collector unknown, NHM. [27°28'S 153°01'E ±10 km]
Other records: Brisbane area (QM specimens).  


Heterocladosoma bifalcatum (Silvestri, 1898)

Heterocladosoma bifalcatum
Eustrongylosoma bifalcatum Silvestri 1898: 231 (first description)
   Figs 13, 14 (p. 231)
Australisoma bifalcatum Brölemann 1913: 89 (in list of Australian species, as Eustrongylosoma bifalcatum)
   91 (in list of species included in Australiosoma, new combination)
   94 (in species key)
Australiosoma bifalcatum Attems 1914: 225 (in species catalogue)
Australiosoma bifalcatum Chamberlin 1920a: 104 (in species catalogue)
Australiosoma bifalcatum Attems 1937: 234 (in species key)
   237 (redescription)
   Fig. 295 (p. 237)
Australiosoma bifalcatus Attems 1944: 249 (distinguished from Australiosoma castaneum, i.e. Oncocladosoma castaneum castaneum)
Heterocladosoma bifalcatum Jeekel 1968: 17 (in list of Australian species, as Australiosoma bifalcatum)
   25 (in family revision, new combination)
   144 (as Eustrongylosoma bifalcatum, designated type species of Heterocladosoma)
   Fig. 2 (p. 145)
Heterocladosoma bifalcatum Jeekel 1981: 47, 49 (in list of species recorded from eastern Australia)
Heterocladosoma bifalcatum Jeekel 1985a: 19 (as Eustrongylosoma bifalcatum, cited as type species of Heterocladosoma)
   22 (in taxonomic discussion)
Heterocladosoma bifalcatum Jeekel 1987: 12 (in list of species recorded from Qld)
   24 (in species key, and redescription)
   Figs 5-8 (p. 26)
Eustrongylosoma bifalcatum Shelley et al. 2000: 105 (in genus catalogue, as type of Heterocladosoma)
Heterocladosoma bifalcatum Rowe and Sierwald 2006: 541 (as Eustrongylosoma bifalcatum, type species of Heterocladosoma; as Australiosoma bifalcatum, in synonymy; redescription)
   Figs 4 (p. 532), 10 (p. 535), 26, 27 (p. 545), 38, 39 (p. 548), 47, 48 (p. 550)
 
Types: At least one male, Cairns, Qld, December 1891, [Giovanni] Podenzana, MSNF.
Other records: Jeekel (1987: 28) believes the Cairns locality is in error and recorded it from Colosseum, Qld (Jeekel 1987: 24). This species is not only abundant in the Brisbane area but is established as far south as Sydney (AM, QM, WAM specimens; Rowe and Sierwald [2006: 541]). It seems to be spreading, probably from southeastern Qld.  


Heterocladosoma galaxias Jeekel, 1985

Heterocladosoma galaxias
Heterocladosoma galaxias Jeekel 1985a: 22 (first description)
   Fig. 2 (p. 23)
Heterocladosoma galaxias Jeekel 1987: 24 (in species key)
Heterocladosoma galaxias Jeekel 2002c: 77 (in list of species recorded from SA)
 
Holotype: Male, Gammon Plateau, SA, 19 September 1956, G.F. Gross, SAM OM1336. [30°32'S 138°33'E ±50 km!!]
Paratypes: None.
Other records: None.  


Heterocladosoma hamuligerum (Verhoeff, 1924)

Heterocladosoma hamuligerum
Australiosoma hamuligerum Verhoeff 1924: 13 (in taxonomic discussion)
   24 (in species key)
   26 (first description)
   Fig. 15, pl. 1
Australiosoma hamuligerum Verhoeff 1928: 88 (in taxonomic discussion)
Australiosoma hamuligerum Attems 1937: 234 (in species key)
   236 (redescription)
Heterocladosoma hamuligerum Jeekel 1968: 17 (in list of Australian species, as Australiosoma hamuligerum)
   25 (in family revision, new combination)
   144 (as Australiosoma hamuligerum, included in new genus Heterocladosoma)
Heterocladosoma hamuligerum Jeekel 1981: 49 (in list of species recorded from eastern Australia)
Heterocladosoma hamuligerum Jeekel 1985a: 22 (in taxonomic discussion)
Heterocladosoma hamuligerum Jeekel 1987: 12 (in list of species recorded from Qld)
   24 (in species key)
   32 (redescription)
   Figs 14, 15 (p. 36)
 
Holotype: Male, Blackall Range, Qld, 19 September 1911, E. Mjöberg, possibly in NHRM. [26°37'S 152°52'S ±5 km]
Paratypes: None.
Other records: Other localities in southeastern Qld (AM, MV, QM specimens).  


Heterocladosoma trabeatum Jeekel, 1987

Heterocladosoma trabeatum
Heterocladosoma trabeatum Jeekel 1987: 12 (in list of species recorded from Qld)
   24 (in species key)
   28 (first description)
   Fig. 10 (p. 30)
 
Holotype: Male, 'Vicinity of Priceden' [?Brisbane], Qld, Eucalyptus forest, under Eucalyptus bark, 28 February 1977, G.F. Kurceva, ZMMU.
Paratypes: None.
Other records: Other localities in central eastern Qld (MV, QM specimens).  


Heterocladosoma transversetaeniatum perarmatum Jeekel, 1987

Heterocladosoma transversetaeniatum perarmatum
Heterocladosoma transversetaeniatum perarmatum Jeekel 1987: 12 (in list of species recorded from Qld)
   31 (first description)
   Figs. 12, 13 (p.33)
 
Holotype: Male, Rockhampton, Qld, 15 November 1924, C. Barrett, AMNH. [23°22'S 150°32'S ±5 km]
Paratypes: 2 males, 1 female, details as for holotype, AMNH.
Other records: None.  


Heterocladosoma transversetaeniatum transversetaeniatum (L. Koch, 1867)

Heterocladosoma transversetaeniatum transversetaeniatum
Strongylosoma transverse-taeniatum L. Koch 1867: 246 [From secondary source, reference not yet checked]
Polydesmus (Strongylosoma) transversetaeniatum Karsch 1881b: 44 (in species list)
   Fig. 23, pl. 3
Strongylosoma transversetaeniatum Attems 1898b: 281 (in species key)
   306 (redescription)
   Figs 18, 19, pl. 1
Australiosoma transverse-taeniatum Brölemann 1913: 89 (in list of Australian species, as Polydesmus (Strongylosoma) transverse-taeniatum)
   91 (in list of species included in Australiosoma, new combination)
   94 (in species key)
Australiosoma transversetaeniatum Brölemann 1916: 539 (in taxonomic discussion)
Australiosoma transversetaeniatum Attems 1914: 225 (in species catalogue)
Australiosoma transversetaeniatum Chamberlin 1920a: 104 (in species catalogue)
Australiosoma transversetaeniatum Attems 1937: 234 (in species key)
   237 (redescription)
   Fig. 296 (p. 238)
Heterocladosoma transversetaeniatum Jeekel 1968: 17 (in list of Australian species, as Australiosoma transversetaeniatum)
   25 (in family revision, new combination)
   144 (as Strongylosoma transversetaeniatum, included in new genus Heterocladosoma)
Heterocladosoma transversetaeniatum Jeekel 1981: 49 (in list of species recorded from eastern Australia)
Heterocladosoma transversetaeniatum Jeekel 1985a: 22 (in taxonomic discussion)
Heterocladosoma transversetaeniatum Jeekel 1987: 12 (in list of species recorded from Qld)
   31 (in taxonomic discussion)
 
Types: The types situation is unclear. Karsch (1881b: 44) says ZMB holds specimens from Cape York (collected by 'Salmin') and Sydney (collected by 'Daemel'). Jeekel (1987) illustrates the right gonopod of 'the male holotype in the Hamburg Museum' (p. 31), from Brisbane, Qld, as drawn by R.L. Hoffman.
Other records: Many specimens (closest to this subspecies) from southeastern Qld in AM and QM.  


Heterocladosoma zebratum Jeekel, 1985

Heterocladosoma zebratum
Heterocladosoma zebratum Jeekel 1985a: 20 (first description)
   Fig. 1 (p. 21)
Heterocladosoma zebratum Jeekel 1987: 24 (in species key)
Heterocladosoma zebratum Jeekel 2002c: 65 (partial redescription)
   77 (in list of species recorded from SA)
   Fig. 4 (p. 62)
 
Holotype: Male, Coralbignie to Buckleboo, SA, 15-16 March 1950, G.F. Gross, SAM, sample missing. [32°46'S 136°16'S ±5 km]
Paratypes:
- 3 females, details as for holotype, SAM OM1327, 1328.
- 1 male, N of Kokotha, SA, 11 June 1956, G.F. Gross, SAM, sample missing. [31°15'S 135°14'S ±10 km]
- 1 male, Nonning, Gawler Ranges, SA, burnt out of spinifex, 17 June 1956, G.F. Gross, 17 June 1956, SAM OM1392. [32°31'S 136°29'S ±10 km]
Other records: Elsewhere in SA (SAM specimens, Jeekel 2002c: 65; WAM specimens).